Answer:
A
P
E
R

We have spotted APER (4 Letters) a total of 161 times in our database. Below you may see the clues associated with APER and also when and where was it last seen:

Definition

• One who apes.

Referring crossword puzzle clues

Usage among publishers:

  • New York Times: Apr 30, 2024
  • Newsday: Jan 2, 2024
  • Newsday: Aug 20, 2023
  • Newsday: Jul 16, 2023
  • Eugene Sheffer: Jul 3, 2023
  • Wall Street Journal: Apr 29, 2023
  • Newsday: Mar 23, 2023